背景1：In recent years, small RNA movement has been both hypothesized and shown to be an integral part of the epigenetic DNA methylation reprogramming occurring during plant reproduction. 

背景2：It was suggested that the release of epigenetic silencing in accessory cell types or tissues is necessary to reinforce epigenetic silencing in the gametes (egg cell and sperm cells), which would in turn ensure the genomic stability of the next generation plant. In Arabidopsis thaliana, small RNA (sRNA) movement was indeed shown to occur during male gametogenesis. 

提出问题：However, the situation within the female gametophyte and in early seed development is mostly unknown. 

结果：Here, we show that small RNAs can induce non-cell-autonomous silencing from the central cell toward the egg cell but also from the synergids to the egg cell and central cell. 

结论：Our data show that in addition to the movement of sRNAs during pollen development, hairpin RNAs can have non-cell-autonomous effects in the female gametes.

近年来，小RNA移动被认为参与了植物生殖过程中表观的DNA甲基化重编程。学者认为，辅助细胞类型或组织中表观遗传沉默的释放对于增强配子（卵细胞和精细胞）中的表观遗传沉默是必要的，这会确保下一世代植物的基因组稳定性。在拟南芥中，确实发现在雄配子发生时，发生了小RNA的移动。但是，雌配子发生以及早期种子发育过程中是否发生小RNA的移动还未可知。本文中，作者发现小RNA从中央细胞到卵细胞以及从助细胞到卵细胞和中央细胞的移动，会诱导非细胞自主性的沉默。作者的实验结果表明除了在花粉发育过程中会发生小RNA的移动，发夹RNA在雌配子体中具有非细胞自主性的效应。